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Several lines of evidence indicate that this form of t-LTD is presynaptic: (i) t-LTD is blocked when presynaptic NMDA receptors are blocked by internal MK-801 in recordings from pairs of synaptically-connected L4 and L2/3 cells (Rodríguez-Moreno and Paulsen, 2008 ), (ii) an increase in paired-pulse ratio is observed after a t-LTD protocol (Bender et al., 2006 ), and (iii) coefficient of variation (CV) analysis is consistent with presynaptic expression (Rodríguez-Moreno and Paulsen, 2008 )
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